Beloved Galápagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died at about 141, the zoo says, adding that "caring for such a remarkable tortoise was a privilege." ...

The California zoo invited fans of the late Galápagos tortoise to enjoy one of her favorite snacks, a "generous, fruit-filled salad," in her honor ...

No one knows exactly when Gramma was born. But if the estimated birth year of 1884 is accurate, Chester Arthur occupied the ...

An occasional series where Mongabay founder Rhett Ayers Butler shares analysis, perspectives and story summaries. She moved slowly, as if time were something best savored. Visitors leaned over ...

Giant tortoises are some of the long-lived species on the planet. Here’s what biologists now understand about one of nature’s ...

Four endangered baby tortoises have made their highly anticipated public debut at the Philadelphia Zoo as the first hatchlings of their 100-year-old mother, Mommy. Mommy, a western Santa Cruz ...

PHILADELPHIA, Pa. (AP) — A pair of critically endangered, nearly 100-year-old Galapagos tortoises at the Philadelphia Zoo have become first-time parents. In an announcement Friday, the zoo said it is ...

Goliath the tortoise had a big shellebration Sunday as he marked his first Father's Day. He also turned 135 at Zoo Miami. His hatchling was born on June 4 after incubating for about four months, ...

LOS ANGELES (AP) — After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has ...